@import url('https://fonts.googleapis.com/css2?family=Fraunces:opsz,wght@9..144,300;9..144,450;9..144,600;9..144,700&family=Source+Sans+3:wght@400;500;600&family=Space+Mono:wght@400;700&display=swap');

:root{
  --ink:#201B18;
  --paper:#EEF0E7;
  --paper-warm:#F7F5EE;
  --plum:#4A2440;
  --plum-deep:#33172D;
  --coral:#C8637A;
  --sage:#6E7F5C;
  --line: rgba(32,27,24,0.14);
}

*{box-sizing:border-box;}
html{scroll-behavior:smooth;}
body{
  margin:0;
  background:var(--paper);
  color:var(--ink);
  font-family:'Source Sans 3', sans-serif;
  font-size:17px;
  line-height:1.6;
}
h1,h2,h3,h4{
  font-family:'Fraunces', serif;
  margin:0;
  font-weight:600;
  letter-spacing:-0.01em;
}
a{color:inherit;}
img{max-width:100%; display:block;}
.mono{
  font-family:'Space Mono', monospace;
  letter-spacing:0.02em;
}
.species{font-style:italic;}
.wrap{
  max-width:1180px;
  margin:0 auto;
  padding:0 40px;
}

/* ---------- NAV ---------- */
nav{
  position:sticky; top:0; z-index:50;
  background:rgba(238,240,231,0.92);
  backdrop-filter:blur(6px);
  border-bottom:1px solid var(--line);
}
nav .wrap{
  display:flex; align-items:center; justify-content:space-between;
  padding-top:18px; padding-bottom:18px;
}
.brand{
  font-family:'Fraunces', serif;
  font-size:19px;
  font-weight:600;
  text-decoration:none;
}
.brand span{color:var(--coral);}
.navlinks{display:flex; gap:34px; list-style:none; margin:0; padding:0;}
.navlinks a{
  text-decoration:none;
  font-size:14px;
  font-weight:600;
  text-transform:uppercase;
  letter-spacing:0.08em;
  color:var(--ink);
  opacity:0.75;
  transition:opacity 0.2s;
}
.navlinks a:hover, .navlinks a.active{opacity:1;}
.navlinks a.active{color:var(--coral); opacity:1;}

/* ---------- HERO (homepage) ---------- */
.hero{
  background:radial-gradient(120% 140% at 15% -10%, var(--plum) 0%, var(--plum-deep) 55%, #221022 100%);
  color:var(--paper-warm);
  position:relative;
  overflow:hidden;
}
.hero .wrap{
  padding-top:90px;
  padding-bottom:70px;
  display:grid;
  grid-template-columns: 1.15fr 0.85fr;
  gap:40px;
  align-items:center;
  position:relative;
  z-index:2;
}
.eyebrow{
  font-family:'Space Mono', monospace;
  font-size:13px;
  letter-spacing:0.12em;
  text-transform:uppercase;
  color:var(--coral);
  margin-bottom:18px;
  display:block;
}
.hero h1{
  font-size:clamp(40px, 6vw, 64px);
  line-height:1.03;
  color:#fff;
}
.hero h1 em{
  font-style:italic;
  color:var(--coral);
  font-weight:600;
}
.hero p.lede{
  margin-top:22px;
  font-size:18px;
  max-width:46ch;
  color:rgba(247,245,238,0.82);
}
.hero-ctas{margin-top:32px; display:flex; gap:16px; flex-wrap:wrap;}
.btn{
  display:inline-block;
  padding:12px 22px;
  border-radius:2px;
  font-size:14px;
  font-weight:600;
  text-decoration:none;
  letter-spacing:0.02em;
  border:1px solid transparent;
  cursor:pointer;
}
.btn-solid{background:var(--coral); color:var(--plum-deep);}
.btn-solid:hover{background:#d67d92;}
.btn-outline{border-color:rgba(247,245,238,0.4); color:var(--paper-warm);}
.btn-outline:hover{border-color:rgba(247,245,238,0.8);}

.hero-art{position:relative; display:flex; flex-direction:column; justify-content:center; align-items:center; text-align:center;}
.hero-art svg{width:100%; max-width:320px; height:auto;}
.hero-art img{
  width:100%;
  max-width:340px;
  aspect-ratio:8/9;
  object-fit:cover;
  border-radius:2px;
  display:block;
}
.hero-art .mono{margin-top:12px; font-size:13px;}
.hero-photo-placeholder{
  width:100%; max-width:320px; aspect-ratio:8/9;
  border:1px dashed rgba(247,245,238,0.35);
  border-radius:4px;
  display:flex; flex-direction:column; align-items:center; justify-content:center; gap:8px;
  color:rgba(247,245,238,0.55);
  text-align:center;
  padding:20px;
}
.hero-photo-placeholder .mono{font-size:13px;}

/* ---------- PAGE HERO (interior pages) ---------- */
.page-hero{
  background:radial-gradient(120% 140% at 15% -10%, var(--plum) 0%, var(--plum-deep) 55%, #221022 100%);
  color:var(--paper-warm);
  padding:64px 0 56px;
}
.page-hero h1{font-size:clamp(32px, 5vw, 46px); color:#fff;}
.page-hero p.lede{margin-top:16px; max-width:60ch; font-size:17px; color:rgba(247,245,238,0.82);}

/* ---------- SECTION SCAFFOLD ---------- */
section{padding:86px 0;}
.section-head{
  display:flex; justify-content:space-between; align-items:flex-end;
  margin-bottom:48px; gap:24px; flex-wrap:wrap;
  border-bottom:1px solid var(--line);
  padding-bottom:22px;
}
.section-head .tag{
  font-family:'Space Mono', monospace;
  font-size:13px;
  letter-spacing:0.1em;
  text-transform:uppercase;
  color:var(--sage);
}
.section-head h2{font-size:32px;}

/* ---------- RESEARCH GRID (homepage teaser + research page) ---------- */
.research-grid{
  display:grid;
  grid-template-columns:repeat(3, 1fr);
  gap:0;
  border-top:1px solid var(--line);
}
.research-col{
  padding:36px 32px 36px 0;
  border-right:1px solid var(--line);
}
.research-col:last-child{border-right:none; padding-right:0;}
.research-col:not(:first-child){padding-left:32px;}
.research-col .num{
  font-family:'Space Mono', monospace;
  font-size:13px;
  color:var(--coral);
  display:block;
  margin-bottom:14px;
}
.research-col h3{font-size:22px; margin-bottom:12px;}
.research-col p{font-size:15.5px; color:rgba(32,27,24,0.78); margin:0;}

/* ---------- RESEARCH PAGE — long-form entries ---------- */
.research-entry{
  display:grid;
  grid-template-columns:200px 1fr;
  gap:36px;
  padding:40px 0;
  border-bottom:1px solid var(--line);
}
.research-entry:first-child{padding-top:0;}
.research-entry .num{
  font-family:'Space Mono', monospace;
  font-size:14px;
  color:var(--coral);
}
.research-entry h3{font-size:26px; margin-bottom:14px;}
.research-entry p{font-size:16px; color:rgba(32,27,24,0.8); margin:0 0 12px;}
.research-entry p:last-child{margin-bottom:0;}
.research-entry ul{
  margin:0 0 12px;
  padding-left:22px;
  font-size:16px;
  color:rgba(32,27,24,0.8);
  line-height:1.6;
}
.research-entry ul li{margin-bottom:6px;}
.research-entry ul li:last-child{margin-bottom:0;}
.research-figure{margin-top:24px; max-width:100%;}
.research-figure .figure-placeholder{
  width:100%; aspect-ratio:16/10;
  border:1px dashed var(--line);
  border-radius:4px;
  display:flex; align-items:center; justify-content:center;
  color:rgba(32,27,24,0.45);
  font-family:'Space Mono', monospace;
  font-size:12.5px;
  text-align:center;
  padding:18px;
}
.research-figure img{width:100%; height:auto; border-radius:4px; display:block;}
.research-figure figcaption{margin-top:8px; font-family:'Space Mono', monospace; font-size:12.5px; color:rgba(32,27,24,0.55); line-height:1.5;}

.research-figure-grid{
  display:grid;
  grid-template-columns:repeat(2, 1fr);
  gap:20px;
  margin-top:24px;
  max-width:100%;
}
.research-figure-grid .research-figure{margin-top:0;}
.research-figure-grid .research-figure img{aspect-ratio:4/3; object-fit:cover;}

.research-figure-grid.stack-right{
  grid-template-rows:repeat(2, 1fr);
}
.research-figure-grid.stack-right .research-figure{
  height:100%;
  display:flex;
  flex-direction:column;
}
.research-figure-grid.stack-right .research-figure img{
  flex:1;
  min-height:0;
  aspect-ratio:auto;
}
.research-figure-grid.stack-right .research-figure:nth-child(1){grid-column:1; grid-row:1;}
.research-figure-grid.stack-right .research-figure:nth-child(2){grid-column:1; grid-row:2;}
.research-figure-grid.stack-right .research-figure:nth-child(3){grid-column:2; grid-row:1 / span 2;}
@media (max-width:640px){
  .research-figure-grid.stack-right{grid-template-rows:none;}
  .research-figure-grid.stack-right .research-figure:nth-child(3){grid-column:1; grid-row:auto;}
}
@media (max-width:640px){
  .research-figure-grid{grid-template-columns:1fr;}
}

/* ---------- GALLERY STRIP (horizontal scroll) ---------- */
.gallery{
  display:flex;
  gap:2px;
  background:var(--ink);
  overflow-x:auto;
  scroll-snap-type:x mandatory;
  -webkit-overflow-scrolling:touch;
  scrollbar-width:thin;
  scrollbar-color:var(--coral) var(--ink);
}
.gallery::-webkit-scrollbar{height:8px;}
.gallery::-webkit-scrollbar-track{background:var(--ink);}
.gallery::-webkit-scrollbar-thumb{background:var(--coral); border-radius:4px;}
.gallery .gallery-item{
  flex:0 0 auto;
  width:260px;
  aspect-ratio:1/1;
  scroll-snap-align:start;
  position:relative;
  overflow:hidden;
  background:linear-gradient(135deg, var(--sage), var(--plum));
  opacity:0.85;
}
.gallery .gallery-item img{
  width:100%; height:100%;
  object-fit:cover;
  display:block;
}
.gallery .gallery-item:nth-child(2n){background:linear-gradient(135deg, var(--coral), var(--plum-deep));}
.gallery .gallery-item:nth-child(3n){background:linear-gradient(135deg, var(--plum), var(--sage));}
.gallery-caption{
  text-align:center;
  font-family:'Space Mono', monospace;
  font-size:12.5px;
  color:rgba(32,27,24,0.55);
  padding:12px 0 0;
  letter-spacing:0.04em;
}
.gallery.auto-scrolling{scroll-snap-type:none; scroll-behavior:auto;}

/* ---------- FUTURE-FEATURE PLACEHOLDER (e.g. iNaturalist feed) ---------- */
.feature-placeholder{
  border:1px dashed var(--line);
  border-radius:4px;
  padding:34px;
  text-align:center;
  color:rgba(32,27,24,0.55);
  font-size:14.5px;
}
.feature-placeholder .mono{display:block; margin-bottom:8px; color:var(--sage);}

/* ---------- PEOPLE (homepage strip) ---------- */
.people-strip{display:flex; gap:28px; overflow-x:auto; padding-bottom:8px;}
.person-card{min-width:190px; flex:0 0 auto;}
.person-card-link{text-decoration:none; color:inherit; display:block; transition:opacity 0.2s;}
.person-card-link:hover{opacity:0.75;}
.person-card{scroll-margin-top:90px;}
.person-photo{width:190px; height:230px; background:linear-gradient(160deg, var(--plum) 0%, var(--sage) 100%); margin-bottom:14px; opacity:0.9; overflow:hidden;}
.person-photo img{width:100%; height:100%; object-fit:cover; display:block;}
.person-card h4{font-family:'Fraunces', serif; font-size:17px; font-weight:600; margin:0 0 4px;}
.person-card .role{font-family:'Space Mono', monospace; font-size:12px; color:var(--sage); text-transform:uppercase; letter-spacing:0.06em;}

/* ---------- PEOPLE PAGE — full grid ---------- */
.people-grid{
  display:grid;
  grid-template-columns:repeat(auto-fill, minmax(220px, 1fr));
  gap:36px 28px;
}
.people-grid .person-photo{width:100%; height:260px;}
.people-grid .person-card{min-width:0;}
.person-card .links{margin-top:8px; display:flex; gap:12px; font-size:13px;}
.person-card .links a{color:var(--sage); text-decoration:none;}
.person-card .links a:hover{color:var(--coral);}
.person-card .bio{font-size:14.5px; color:rgba(32,27,24,0.72); margin-top:10px;}

.recruiting{
  margin-top:30px;
  padding:22px 26px;
  background:var(--paper-warm);
  border-left:3px solid var(--coral);
  font-size:15.5px;
}
.recruiting strong{font-family:'Fraunces', serif; font-weight:600;}

/* ---------- PUBLICATIONS ---------- */
.pub-filter{display:flex; gap:10px; margin-bottom:32px; flex-wrap:wrap;}
.pub-filter button{
  font-family:'Space Mono', monospace;
  font-size:12.5px;
  text-transform:uppercase;
  letter-spacing:0.05em;
  padding:8px 16px;
  background:transparent;
  border:1px solid var(--line);
  border-radius:20px;
  cursor:pointer;
  color:var(--ink);
}
.pub-filter button.active{background:var(--plum); color:#fff; border-color:var(--plum);}
.pub-list{list-style:none; margin:0; padding:0;}
.pub-list li{display:grid; grid-template-columns:70px 1fr; gap:24px; padding:20px 0; border-bottom:1px solid var(--line);}
.pub-list li:first-child{padding-top:0;}
.pub-year{font-family:'Space Mono', monospace; font-size:14px; color:var(--coral); padding-top:2px;}
.pub-title{font-size:16px; margin-bottom:4px;}
.pub-meta{font-size:14px; color:rgba(32,27,24,0.6);}
.pub-links{margin-top:6px; display:flex; gap:14px;}
.pub-links a{font-size:13px; font-family:'Space Mono', monospace; color:var(--sage); text-decoration:none;}
.pub-links a:hover{color:var(--coral);}

/* ---------- CONTACT PAGE ---------- */
.contact-grid{display:grid; grid-template-columns:1fr 1fr; gap:60px;}
.contact-block h3{font-size:20px; margin-bottom:14px;}
.contact-block p{font-size:15.5px; color:rgba(32,27,24,0.78);}
.contact-list{list-style:none; margin:0; padding:0;}
.contact-list li{padding:10px 0; border-bottom:1px solid var(--line); display:flex; justify-content:space-between; font-size:15px;}
.contact-list li a{color:var(--sage); text-decoration:none;}
.contact-list li a:hover{color:var(--coral);}

/* ---------- FOOTER ---------- */
footer{background:var(--plum-deep); color:rgba(247,245,238,0.75); padding:56px 0 34px;}
footer .wrap{display:flex; justify-content:space-between; align-items:flex-start; flex-wrap:wrap; gap:24px;}
footer h3{color:#fff; font-size:20px; margin-bottom:10px;}
footer a{text-decoration:none; color:rgba(247,245,238,0.85);}
footer a:hover{color:var(--coral);}
.foot-links{display:flex; gap:22px; list-style:none; padding:0; margin:0; font-size:14px;}
.copyright{margin-top:40px; padding-top:20px; border-top:1px solid rgba(247,245,238,0.15); font-family:'Space Mono', monospace; font-size:12.5px; color:rgba(247,245,238,0.5);}

@media (max-width:860px){
  .hero .wrap{grid-template-columns:1fr; padding-top:60px;}
  .hero-art{order:-1; margin-bottom:12px;}
  .research-grid{grid-template-columns:1fr;}
  .research-col{border-right:none; padding:28px 0 !important; border-bottom:1px solid var(--line);}
  .research-entry{grid-template-columns:1fr; gap:10px;}
  .gallery{grid-template-columns:repeat(3,1fr);}
  .navlinks{display:none;}
  section{padding:56px 0;}
  .wrap{padding:0 24px;}
  .contact-grid{grid-template-columns:1fr; gap:36px;}
}